# Tinbergen's four questions > Nikolaas Tinbergen의 네 가지 질문. 의 네 가지 질문. - Behavioural adaptive functions - Phylogenetic history; and the proximate explanations - Underlying physiological mechanisms - Ontogenetic/developmental history. ## Example [Visual perception](https://wiki.g15e.com/pages/Visual%20perception.txt): - Function: To find food and avoid danger. - Phylogeny: The vertebrate eye initially developed with a blind spot, but the lack of adaptive intermediate forms prevented the loss of the blind spot. - Causation: The lens of the eye focuses light on the retina. - Development: Neurons need the stimulation of light to wire the eye to the brain